- David Y. Cooper
Lusk, Graham (15 February 1866–18 July 1932), physiologist and biochemist, was born in Bridgeport, Connecticut, the son of William Thompson Lusk, an obstetrician, and Mary Hartwell Chittenden. Having impaired hearing, young Lusk followed his father’s advice not to become a physician and instead studied chemistry at the Columbia School of Mines in New York, graduating with a Ph.B. in 1887. To study the biological sciences, Lusk traveled to Europe; in the next few years he worked first in Leipzig under the famous physiologist Carl Ludwig and then in Munich under the physiological chemist Carl Voit. In 1891 he received his Ph.D. from the University of Munich and returned to America full of enthusiasm for the Voit-Rubner doctrines in nutrition, which held that the energy derived from the metabolism of the three groups of foodstuffs—carbohydrate, fat, and protein—was exchangeable in the body in accordance with caloric equivalence. Voit and Rubner also stated that the metabolic rate was related to the body surface area of the individual....