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Keeler, Clyde Edgar (11 April 1900–22 April 1994), biologist, educator, and cultural historian, was born in Marion, Ohio, the son of Anthony Sylvester Keeler, a watchmaker and teacher, and Amanda Jane Dumm Keeler, a teacher. Growing up in Marion, with nearby farmlands, Keeler had early opportunities—on his milk and paper routes—to observe nature, and he attributed the launching of his biomedical career to childhood observations of field mice. Keeler graduated from Denison University (Granville, Ohio) in 1923 with a zoology major and enough credits for a master’s degree; he lacked only the research component, which he completed in 1925 at Harvard. Cited as “the school artist” in the yearbook, he was Phi Beta Kappa, president of the Zoology Club, and captain of the cross country team. He was also a member of the Student Army Training Corps (for World War I) and, after the war, the Reserve Officers Training Corps; he eventually rose to the rank of major in the U.S. Army Officers Reserve Corps....